This expression comes down from the days of horse drawn wagons and coaches where on journeys through unsafe areas someone would sit next to the driver holding a weapon (sometimes literally a shotgun). When someone "calls shotgun" they are claiming the front passenger seat. There are rules about when and how you can "call shotgun". The most common rule is that you must be within sight of the car. "Calling shotgun" is an informal practice whereby friends can decide who will sit where without an argument.
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